About the Authors

Beth K Thielen

Affiliation Department of Pathobiology, University of Washington, Seattle, Washington, United States of America

Kevin C Klein

Affiliation Department of Pathobiology, University of Washington, Seattle, Washington, United States of America

Lorne W Walker

Affiliation Department of Pathobiology, University of Washington, Seattle, Washington, United States of America

Mary Rieck

Affiliation Translational Research Program, Benaroya Research Institute at Virginia Mason, Seattle, Washington, United States of America

Jane H Buckner

Affiliation Translational Research Program, Benaroya Research Institute at Virginia Mason, Seattle, Washington, United States of America

Garrett W Tomblingson

Affiliation Department of Pathobiology, University of Washington, Seattle, Washington, United States of America

Jaisri R Lingappa

To whom correspondence should be addressed. E-mail: jais@u.washington.edu

Affiliations Department of Pathobiology, University of Washington, Seattle, Washington, United States of America , Department of Medicine, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

BKT, KCK, LWW, and JRL conceived and designed the experiments. BKT and MR performed the experiments. BKT analyzed the data. JHB and GWT contributed reagents, materials, and/or analysis tools. BKT, KCK, and JRL wrote the paper.