About the Authors

Christopher J Johnson

Affiliations Program in Cellular and Molecular Biology, University of Wisconsin Madison, Madison, Wisconsin, United States of America , Department of Animal Health and Biomedical Sciences, School of Veterinary Medicine, University of Wisconsin Madison, Madison, Wisconsin, United States of America

Kristen E Phillips

Affiliation Molecular and Environmental Toxicology Center, University of Wisconsin Madison, Madison, Wisconsin, United States of America

Peter T Schramm

Affiliation Molecular and Environmental Toxicology Center, University of Wisconsin Madison, Madison, Wisconsin, United States of America

Debbie McKenzie

Affiliation Department of Animal Health and Biomedical Sciences, School of Veterinary Medicine, University of Wisconsin Madison, Madison, Wisconsin, United States of America

Judd M Aiken

Affiliations Program in Cellular and Molecular Biology, University of Wisconsin Madison, Madison, Wisconsin, United States of America , Department of Animal Health and Biomedical Sciences, School of Veterinary Medicine, University of Wisconsin Madison, Madison, Wisconsin, United States of America

Joel A Pedersen

To whom correspondence should be addressed. E-mail: joelpedersen@wisc.edu

Affiliations Molecular and Environmental Toxicology Center, University of Wisconsin Madison, Madison, Wisconsin, United States of America , Department of Soil Science, University of Wisconsin Madison, Madison, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

CJJ, DM, JMA, and JAP conceived and designed the experiments. CJJ, KEP, and PTS performed the experiments. CJJ, KEP, PTS, DM, JMA, and JAP analyzed the data. JMA and JAP contributed reagents/materials/analysis tools. CJJ, DM, JMA, and JAP wrote the paper.