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Contributed equally to this work with:
Michelle L. Verant,
Justin G. Boyles
Department of Pathobiological Sciences, University of Wisconsin, Madison, Wisconsin, United States of America
Department of Ecology and Evolutionary Biology, University of Tennessee, Knoxville, Tennessee, United States of America
Current address: Cooperative Wildlife Research Laboratory, Department of Zoology, Southern Illinois University, Carbondale, Illinois, United States of America
Department of Computer Science, University of Chicago, Chicago, Illinois, United States of America
Leibniz Institute for Zoo and Wildlife Research, Berlin, Germany
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US Geological Survey – National Wildlife Health Center, Madison, Wisconsin, United States of America
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