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Contributed equally to this work with:
Jeannette E. Loram
Interdisciplinary Program in Genetics, University of Iowa, Iowa City, Iowa, United States of America
Bermuda Institute of Ocean Sciences, St. George's, Bermuda
Ecology and Evolutionary Biology Department, University of Arizona, Tucson, Arizona, United States of America
Woods Hole Oceanographic Institution, Woods Hole, Massachusetts, United States of America
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Interdisciplinary Program in Genetics, University of Iowa, Iowa City, Iowa, United States of America,
Department of Biology, University of Iowa, Iowa City, Iowa, United States of America
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