Complete Primate Skeleton from the Middle Eocene of Messel in Germany: Morphology and Paleobiology
(A)— Europolemur klatti (Weigelt, 1933), GMH CeIV-3656, left and right mandible with I1–2, C1, P2-M3d and I1–2, C1, P3-M3s. (B)— Europolemur klatti (Weigelt, 1933), GMH LeoI-4233, part of the skull with upper dentition, which is part of the holotype. (C)— Europolemur klatti (Weigelt, 1933), GMH XXXVII-120, fragmentary left mandible with double-rooted P2, P3–4, and heavily worn M1–3. (D)— Europolemur klatti (Weigelt, 1933), GMH XXII-1, right mandibular ramus with P3-M3 and alveoli for a double rooted P2. (E)— Protoadapis ignoratus (Thalmann, 1994), GMH XXII-549, part of type specimen, fragment of right mandible with C1, P3–4, M1, alveoli of P2 and M3 (mirrored). (F)— Protoadapis weigelti Gingerich, 1977, GMH XXII-624, right mandibular ramus with P3-M3, root of a small single-rooted P2 and alveolus of C1, which is isolated (mirrored). (G)— Godinotia neglecta (Thalmann, Haubold & Martin, 1989), holotype (GMH L-2), detail: palate containing M3-P3s and d, and the small unicuspid and one-rooted P2s. Arrows show the position of P2/P2. Geiseltal primates come from Middle Eocene zones MP12 and 13, slightly later in time than those from Messel (MP11). Godinotia neglecta, like Darwinius masillae, is distinguished from Europolemur klatti by the presence of small, straight, single-rooted P2.