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Contributed equally to this work with:
Current address: Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom
Animal Health Trust, Lanwades Park, Kentford, Newmarket, Suffolk, United Kingdom
Institute for Animal Health, Pirbright Laboratory, Pirbright, Surrey, United Kingdom
Current address: Clemson University, Clemson, South Carolina, United States of America
Cambridge Infectious Diseases Consortium, Department of Veterinary Medicine, Cambridge, United Kingdom
Sir Albert Sakzewski Virus Research Centre, University of Queensland, Herston, Queensland, Australia
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Current address: Institute for Animal Health, Pirbright Laboratory, Pirbright, Surrey, United Kingdom
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