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Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan
International Coastal Research Center, Ocean Research Institute, The University of Tokyo, Otsuchi, Iwate, Japan
Graduate School of Fisheries Science, Hokkaido University, Hakodate, Hokkaido, Japan
National Institute of Polar Research, Itabashi, Tokyo, Japan
Centre for Ecology and Hydrology, Bush Estate, Penicuik, Midlothian, United Kingdom
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