About the Authors

Janna R. Willoughby

willoughby@purdue.edu

Affiliations Department of Forestry and Natural Resources, Purdue University, West Lafayette, Indiana, United States of America, Department of Biological Sciences, Purdue University, West Lafayette, Indiana, United States of America

Jamie A. Ivy

Affiliation San Diego Zoo Global Collections Department, San Diego, California, United States of America

Robert C. Lacy

Affiliation Chicago Zoological Society, Brookfield, Illinois, United States of America

Jacqueline M. Doyle

Affiliation Department of Biological Sciences, Towson University, Towson, Maryland, United States of America

J. Andrew DeWoody

Affiliations Department of Forestry and Natural Resources, Purdue University, West Lafayette, Indiana, United States of America, Department of Biological Sciences, Purdue University, West Lafayette, Indiana, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: JRW JAI RCL JAD. Formal analysis: JRW JMD. Funding acquisition: JRW JAI RCL JAD. Investigation: JRW JAI RCL JMD. Resources: JRW JAI RCL JAD. Software: JRW JMD. Writing – original draft: JRW JAD. Writing – review & editing: JRW JAI RCL JMD JAD.