About the Authors

Nicholas G. Dannemiller

dannemillern@gmail.com

Affiliations Department of Fish, Wildlife, and Conservation Biology, Colorado State University, Fort Collins, Colorado, United States of America, United States Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Colleen T. Webb

Affiliation Department of Biology, Colorado State University, Fort Collins, Colorado, United States of America

Kenneth R. Wilson

Affiliation Department of Fish, Wildlife, and Conservation Biology, Colorado State University, Fort Collins, Colorado, United States of America

Kevin T. Bentler

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Nicole L. Mooers

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Jeremy W. Ellis

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

J. Jeffrey Root

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Alan B. Franklin

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Susan A. Shriner

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: NGD CTW KRW SAS. Formal analysis: NGD SAS. Investigation: NGD KTB NLM JWE JJR. Methodology: NGD CTW SAS. Project administration: NGD SAS. Writing – original draft: NGD CTW SAS. Writing – review & editing: NGD CTW KRW KTB NLM JWE JJR ABF SAS.