About the Authors

Lan-Hai Wei

Contributed equally to this work with: Lan-Hai Wei, Shi Yan

Affiliations MOE Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, China, Institut National des Langues et Civilisations Orientales, Paris, France

Shi Yan

Contributed equally to this work with: Lan-Hai Wei, Shi Yan

Affiliation MOE Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, China

Yik-Ying Teo

Affiliation Saw Swee Hock School of Public Health, National University of Singapore, Singapore, Singapore

Yun-Zhi Huang

Affiliation MOE Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, China

Ling-Xiang Wang

Affiliation MOE Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, China

Ge Yu

Affiliation MOE Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, China

Woei-Yuh Saw

Affiliation Saw Swee Hock School of Public Health, National University of Singapore, Singapore, Singapore

Rick Twee-Hee Ong

Affiliation Saw Swee Hock School of Public Health, National University of Singapore, Singapore, Singapore

Yan Lu

Affiliation Chinese Academy of Sciences and Max Planck Society (CAS-MPG) Partner Institute for Computational Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, China

Chao Zhang

Affiliations Chinese Academy of Sciences and Max Planck Society (CAS-MPG) Partner Institute for Computational Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, China, University of Chinese Academy of Sciences, Beijing, China

Shu-Hua Xu

Affiliations Chinese Academy of Sciences and Max Planck Society (CAS-MPG) Partner Institute for Computational Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, China, University of Chinese Academy of Sciences, Beijing, China, School of Life Science and Technology, ShanghaiTech University, Shanghai, China, Collaborative Innovation Center of Genetics and Development, Shanghai, China

Li Jin

Affiliations MOE Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, China, Collaborative Innovation Center of Genetics and Development, Shanghai, China

Hui Li

LHCA@Fudan.edu.cn

Affiliation MOE Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: LHW HL. Formal analysis: LHW SY YYT YZH LXW WYS RT-HO CZ YL SHX LJ HL. Investigation: GY CZ YL. Resources: HL YYT SHX LJ. Writing – original draft: HL LHW. Writing – review & editing: LHW SY YYT YZH LXW WYS RT-HO CZ YL SHX LJ HL.