About the Authors

Leonie Verhage

Contributed equally to this work with: Leonie Verhage, Edouard I. Severing

Affiliations Laboratory of Molecular Biology, Wageningen University and Research, PB Wageningen, The Netherlands, Bioscience, Wageningen Plant Research, Wageningen University and Research, PB Wageningen, The Netherlands

Edouard I. Severing

Contributed equally to this work with: Leonie Verhage, Edouard I. Severing

Affiliations Bioscience, Wageningen Plant Research, Wageningen University and Research, PB Wageningen, The Netherlands, Max Planck Institute for Plant Breeding Research, Köln, Germany

Johan Bucher

Affiliation Wageningen UR Plant Breeding, Wageningen University and Research, PB Wageningen, The Netherlands

Michiel Lammers

Affiliation Bioscience, Wageningen Plant Research, Wageningen University and Research, PB Wageningen, The Netherlands

Jacqueline Busscher-Lange

Affiliation Bioscience, Wageningen Plant Research, Wageningen University and Research, PB Wageningen, The Netherlands

Guusje Bonnema

Affiliation Wageningen UR Plant Breeding, Wageningen University and Research, PB Wageningen, The Netherlands

Nicole Rodenburg

Affiliation Molecular Plant Physiology, Institute of Environmental Biology, Utrecht University, CH Utrecht, The Netherlands

Marcel C. G. Proveniers

Affiliation Molecular Plant Physiology, Institute of Environmental Biology, Utrecht University, CH Utrecht, The Netherlands

Gerco C. Angenent

Affiliations Laboratory of Molecular Biology, Wageningen University and Research, PB Wageningen, The Netherlands, Bioscience, Wageningen Plant Research, Wageningen University and Research, PB Wageningen, The Netherlands

Richard G. H. Immink

richard.immink@wur.nl

Affiliations Laboratory of Molecular Biology, Wageningen University and Research, PB Wageningen, The Netherlands, Bioscience, Wageningen Plant Research, Wageningen University and Research, PB Wageningen, The Netherlands

Competing Interests

Funding was received from a commercial source: Syngenta, Bejo Seeds B.V. and Rijk Zwaan Breeding B.V. This does not alter our adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: GB MCGP GCA RGHI. Data curation: EIS. Formal analysis: LV EIS NR. Funding acquisition: GB MCGP GCA RGHI. Investigation: LV EIS JBL ML JB NR. Methodology: LV EIS GB MCGP GCA RGHI. Project administration: GCA GB RGHI. Resources: GCA GB MCGP. Software: EIS. Supervision: GB MCGP GCA RGHI. Validation: LV EIS NR MCGP GCA RGHI. Visualization: LV EIS. Writing – original draft: LV EIS. Writing – review & editing: GB GCA RGHI.