About the Authors

Wangxiang Zhang

Contributed equally to this work with: Wangxiang Zhang, Junjun Fan

malus2011@163.com

Affiliations College of Forestry, Nanjing Forestry University, Nanjing, China, Co-Innovation Center for Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing, China, Yangzhou Crabapple Horticulture Company Limited, Yangzhou, China

Junjun Fan

Contributed equally to this work with: Wangxiang Zhang, Junjun Fan

Affiliations College of Forestry, Nanjing Forestry University, Nanjing, China, Co-Innovation Center for Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing, China

Qianqian Tan

Affiliation Suzhou Forestry Station, Suzhou, China

Mingming Zhao

Affiliations College of Forestry, Nanjing Forestry University, Nanjing, China, Co-Innovation Center for Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing, China

Ting Zhou

Affiliations College of Forestry, Nanjing Forestry University, Nanjing, China, Co-Innovation Center for Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing, China

Fuliang Cao

Affiliations College of Forestry, Nanjing Forestry University, Nanjing, China, Co-Innovation Center for Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing, China

Competing Interests

WZ is an employee of Yangzhou Crabapple Horticulture Company Limited. This does not alter our adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: WZ. Formal analysis: JF. Investigation: QT. Methodology: WZ. Supervision: FC. Writing – original draft: WZ JF. Writing – review & editing: MZ TZ.