About the Authors

Kelly A. Williams

williak5@ohio.edu

Affiliation Department of Biological Sciences, Ohio University, Athens, OH 45701, United States of America

Donald B. Miles

Affiliation Department of Biological Sciences, Ohio University, Athens, OH 45701, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: KAW DBM. Data curation: KAW. Formal analysis: KAW. Funding acquisition: KAW. Investigation: KAW. Methodology: KAW. Supervision: DBM. Writing – original draft: KAW. Writing – review & editing: KAW DBM.