About the Authors

Haiyang Zuo

Contributed equally to this work with: Haiyang Zuo, Junfang Zhang, Liuguang Zhang, Xiaoxia Ren

Affiliations The Key Laboratory for Farm Animal Genetic and Utilization of Ministry of Agriculture of China, Institute of Animal Science, Chinese Academy of Agricultural Science, Beijing, 100193, China, JiLin Agriculture University Animal Science and Technology, Changchun, 130118, China

Junfang Zhang

Contributed equally to this work with: Haiyang Zuo, Junfang Zhang, Liuguang Zhang, Xiaoxia Ren

Affiliation The Key Laboratory for Farm Animal Genetic and Utilization of Ministry of Agriculture of China, Institute of Animal Science, Chinese Academy of Agricultural Science, Beijing, 100193, China

Liuguang Zhang

Contributed equally to this work with: Haiyang Zuo, Junfang Zhang, Liuguang Zhang, Xiaoxia Ren

Affiliations The Key Laboratory for Farm Animal Genetic and Utilization of Ministry of Agriculture of China, Institute of Animal Science, Chinese Academy of Agricultural Science, Beijing, 100193, China, HaiKou Mary Hospital Reproductive Medicine Center, Haikou, 570100, China

Xiaoxia Ren

Contributed equally to this work with: Haiyang Zuo, Junfang Zhang, Liuguang Zhang, Xiaoxia Ren

Affiliation The Key Laboratory for Farm Animal Genetic and Utilization of Ministry of Agriculture of China, Institute of Animal Science, Chinese Academy of Agricultural Science, Beijing, 100193, China

Xiaoli Chen

‡ These authors also contributed equally to this work.

Affiliation The Key Laboratory for Farm Animal Genetic and Utilization of Ministry of Agriculture of China, Institute of Animal Science, Chinese Academy of Agricultural Science, Beijing, 100193, China

Haisheng Hao

‡ These authors also contributed equally to this work.

Affiliation The Key Laboratory for Farm Animal Genetic and Utilization of Ministry of Agriculture of China, Institute of Animal Science, Chinese Academy of Agricultural Science, Beijing, 100193, China

Xueming Zhao

‡ These authors also contributed equally to this work.

Affiliation The Key Laboratory for Farm Animal Genetic and Utilization of Ministry of Agriculture of China, Institute of Animal Science, Chinese Academy of Agricultural Science, Beijing, 100193, China

Dong Wang

dwangcn2002@vip.sina.com

Affiliation The Key Laboratory for Farm Animal Genetic and Utilization of Ministry of Agriculture of China, Institute of Animal Science, Chinese Academy of Agricultural Science, Beijing, 100193, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: DW. Data curation: JZ. Formal analysis: LZ JZ. Funding acquisition: DW. Methodology: DW. Project administration: XC HH XZ. Resources: DW. Software: LZ JZ HZ. Supervision: DW. Validation: HZ XR. Writing – original draft: HZ JZ. Writing – review & editing: DW.