About the Authors

Zheguang Lin

Contributed equally to this work with: Zheguang Lin, Paul Page

Affiliation College of Animal Sciences, Zhejiang University, Hangzhou, China

Paul Page

Contributed equally to this work with: Zheguang Lin, Paul Page

Affiliation Agroscope, Swiss Bee Research Center, Bern, Switzerland

Li Li

Affiliation College of Animal Sciences, Zhejiang University, Hangzhou, China

Yao Qin

Affiliation College of Animal Sciences, Zhejiang University, Hangzhou, China

Yingying Zhang

Affiliation College of Animal Sciences, Zhejiang University, Hangzhou, China

Fuliang Hu

Affiliation College of Animal Sciences, Zhejiang University, Hangzhou, China

Peter Neumann

Affiliations Institute of Bee Health, Vetsuisse Faculty, University of Bern, Bern, Switzerland, Bee Protection Laboratory, Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand, Social Insect Research Group, Department of Zoology and Entomology, University of Pretoria, Pretoria, South Africa

Huoqing Zheng

hqzheng@zju.edu.cn

Affiliation College of Animal Sciences, Zhejiang University, Hangzhou, China

Vincent Dietemann

Affiliations Agroscope, Swiss Bee Research Center, Bern, Switzerland, Social Insect Research Group, Department of Zoology and Entomology, University of Pretoria, Pretoria, South Africa

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: VD PN HZ FH. Data curation: ZL PP LL YQ YZ. Formal analysis: ZL PP VD. Funding acquisition: FH HZ VD PN. Investigation: PP ZL LL YQ YZ. Methodology: VD PP ZL LL YQ YZ. Project administration: HZ FH VD PN. Resources: FH HZ VD PN. Supervision: HZ FH VD PN. Validation: VD PN HZ FH. Writing – original draft: ZL PP. Writing – review & editing: VD PN PP HZ ZL.