About the Authors

Anurag Verma

Affiliations Biomedical and Translational Informatics Program, Geisinger Health System, Danville, Pennsylvania, United States of America, Center for Systems Genomics, Department of Biochemistry and Molecular Biology, The Huck Institutes of the Life Sciences, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Anna O. Basile

Affiliation Center for Systems Genomics, Department of Biochemistry and Molecular Biology, The Huck Institutes of the Life Sciences, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Yuki Bradford

Affiliation Biomedical and Translational Informatics Program, Geisinger Health System, Danville, Pennsylvania, United States of America

Helena Kuivaniemi

Affiliations The Sigfried and Janet Weis Center for Research, Geisinger Health System, Danville, Pennsylvania, United States of America, Division of Molecular Biology and Human Genetics, Department of Biomedical Sciences, Faculty of Medicine and Health Sciences, Stellenbosch University, Tygerberg, South Africa

Gerard Tromp

Affiliations The Sigfried and Janet Weis Center for Research, Geisinger Health System, Danville, Pennsylvania, United States of America, Division of Molecular Biology and Human Genetics, Department of Biomedical Sciences, Faculty of Medicine and Health Sciences, Stellenbosch University, Tygerberg, South Africa

David Carey

Affiliation The Sigfried and Janet Weis Center for Research, Geisinger Health System, Danville, Pennsylvania, United States of America

Glenn S. Gerhard

Affiliation Department of Medical Genetics and Molecular Biochemistry, Temple University School of Medicine, Philadelphia, Pennsylvania, United States of America

James E. Crowe Jr

Affiliation The Vanderbilt Vaccine Center, Vanderbilt University, Nashville Tennessee, United States of America

Marylyn D. Ritchie

Affiliations Biomedical and Translational Informatics Program, Geisinger Health System, Danville, Pennsylvania, United States of America, Center for Systems Genomics, Department of Biochemistry and Molecular Biology, The Huck Institutes of the Life Sciences, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Sarah A. Pendergrass

spendergrass@geisinger.edu

Affiliation Biomedical and Translational Informatics Program, Geisinger Health System, Danville, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AV AB HK GT JC MR SP. Performed the experiments: YB. Analyzed the data: AV AB SP. Wrote the paper: AV AB HK DC GG JC MR SP.