About the Authors

Ruth V. Nichols

Affiliations Grimsö Wildlife Research Station, Department of Ecology, Swedish University of Agricultural Sciences, Riddarhyttan, Sweden, Molecular Ecology Group, Department of Wildlife, Fish and Environmental Studies, Swedish University of Agricultural Sciences, Skogsmarksgränd, Umeå, Sweden, Department of Ecology and Evolutionary Biology, University of California Santa Cruz, Santa Cruz, California, United States of America

Mikael Åkesson

mikael.akesson@slu.se

Affiliation Grimsö Wildlife Research Station, Department of Ecology, Swedish University of Agricultural Sciences, Riddarhyttan, Sweden

Petter Kjellander

Affiliation Grimsö Wildlife Research Station, Department of Ecology, Swedish University of Agricultural Sciences, Riddarhyttan, Sweden

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PK MÅ. Performed the experiments: MÅ. Analyzed the data: RN MÅ. Wrote the paper: RN MÅ PK.