About the Authors

Hung-Chang Shen

Contributed equally to this work with: Hung-Chang Shen, Jia-Yi Wei, Sao-Yu Chu, Pei-Chi Chung, Tsai-Chi Hsu

Affiliations Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan, Graduate Institute of Life Sciences, National Defense Medical Center, Taipei, Taiwan

Jia-Yi Wei

Contributed equally to this work with: Hung-Chang Shen, Jia-Yi Wei, Sao-Yu Chu, Pei-Chi Chung, Tsai-Chi Hsu

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Sao-Yu Chu

Contributed equally to this work with: Hung-Chang Shen, Jia-Yi Wei, Sao-Yu Chu, Pei-Chi Chung, Tsai-Chi Hsu

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Pei-Chi Chung

Contributed equally to this work with: Hung-Chang Shen, Jia-Yi Wei, Sao-Yu Chu, Pei-Chi Chung, Tsai-Chi Hsu

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Tsai-Chi Hsu

Contributed equally to this work with: Hung-Chang Shen, Jia-Yi Wei, Sao-Yu Chu, Pei-Chi Chung, Tsai-Chi Hsu

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Hung-Hsiang Yu

samhhyu@gate.sinica.edu.tw

Affiliations Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan, Graduate Institute of Life Sciences, National Defense Medical Center, Taipei, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HCS JYW HHY. Performed the experiments: HCS JYW SYC PCC TCH. Analyzed the data: HCS JYW SYC PCC. Wrote the paper: HHY.