About the Authors

Ian M. McLeod

ianmcleodnz@gmail.com

Affiliations Centre for Tropical Water and Aquatic Ecosystem Research (TropWATER), College of Science and Engineering and ARC Centre of Excellence for Coral Reef Studies, James Cook University, Townsville, Queensland, Australia, Australian Institute of Marine Science and AIMS@JCU, Townsville, Queensland, Australia

Timothy D. Clark

Current address: University of Tasmania and CSIRO Agriculture Flagship, Hobart, Australia

Affiliation Australian Institute of Marine Science and AIMS@JCU, Townsville, Queensland, Australia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: IMM TDC. Performed the experiments: IMM TDC. Analyzed the data: IMM TDC. Contributed reagents/materials/analysis tools: IMM TDC. Wrote the paper: IMM TDC.