About the Authors

Joseph P. Garner

Affiliation Stanford University, Department of Comparative Medicine, and by courtesy, Department of Psychiatry and Behavioral Sciences, Stanford, California, United States of America

Brianna N. Gaskill

Affiliations Charles River, Wilmington, Massachusetts, United States of America, Purdue University Department of Comparative Pathobiology, West Lafayette, Indiana, United States of America

Kathleen R. Pritchett-Corning

pritchettcorning@fas.harvard.edu

Affiliations Charles River, Wilmington, Massachusetts, United States of America, Harvard University Faculty of Arts and Sciences, Office of Animal Resources, Cambridge, Massachusetts, United States of America

Competing Interests

KPC and BNG were employees of Charles River at the time this work was performed and both are currently consultants for the company. KPC is also an academic editor at PLOS One. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JPG KPC. Performed the experiments: BNG KPC. Analyzed the data: BNG JPG. Wrote the paper: KPC BNG JPG.