About the Authors

Claire Heffernan

ch15664@bristol.ac.uk

Affiliation School of Veterinary Sciences, University of Bristol, Langford, Bristol, United Kingdom

Lena Azbel-Jackson

Affiliation Livestock Development Group, University of Reading, Reading, United Kingdom

Joe Brownlie

Affiliation Royal Veterinary College, North Mymms, Hatfield, Herts, United Kingdom

George Gunn

Affiliation Scotland’s Rural College, Aberdeen, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CH. Performed the experiments: CH. Analyzed the data: CH LA-J. Wrote the paper: CH L-AJ GG JB.