About the Authors

Julie Beckstead

Affiliation Department of Biology, Gonzaga University, Spokane, Washington, 99258, United States of America

Susan E. Meyer

semeyer@xmission.com

Affiliation USDA Forest Service, Rocky Mountain Research Station, Shrub Sciences Laboratory, Provo, UT 84606, United States of America

Toby S. Ishizuka

Affiliation Department of Biology, Gonzaga University, Spokane, Washington, 99258, United States of America

Kelsey M. McEvoy

Affiliation Department of Biology, Gonzaga University, Spokane, Washington, 99258, United States of America

Craig E. Coleman

Affiliation Department of Plant and Wildlife Sciences, Brigham Young University, Provo, UT 84602, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SEM JB. Performed the experiments: JB TSI KMM CEC. Analyzed the data: SEM JB TSI KMM CEC. Contributed reagents/materials/analysis tools: CEC. Wrote the paper: SEM JB TSI KMM. Critically reviewed the manuscript: CEC.