About the Authors

Beverly Paigen

Contributed equally to this work with: Beverly Paigen, Karen L. Svenson

Affiliation The Jackson Laboratory, 600 Main Street, Bar Harbor, Maine, 04609, United States of America

Joanne M. Currer

Affiliation The Jackson Laboratory, 600 Main Street, Bar Harbor, Maine, 04609, United States of America

Karen L. Svenson

Contributed equally to this work with: Beverly Paigen, Karen L. Svenson

Karen.Svenson@jax.org

Affiliation The Jackson Laboratory, 600 Main Street, Bar Harbor, Maine, 04609, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KLS BP. Performed the experiments: BP KLS. Analyzed the data: BP KLS. Wrote the paper: BP JMC KLS.