About the Authors

Guilin Li

Contributed equally to this work with: Guilin Li, Hang Zhao

Affiliation State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian, Shandong, 271018, P. R. China

Hang Zhao

Contributed equally to this work with: Guilin Li, Hang Zhao

Affiliation State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian, Shandong, 271018, P. R. China

Hongfang Wang

Affiliation College of Animal Science and Technology, Shandong Agricultural University, Taian, Shandong, 271018, P. R. China

Xulei Guo

Affiliation State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian, Shandong, 271018, P. R. China

Xingqi Guo

Affiliation State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian, Shandong, 271018, P. R. China

Qinghua Sun

qhsun@sdau.edu.cn (QS); bhxu@sdau.edu.cn (BX)

‡ These authors also contributed equally to this work.

Affiliation State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian, Shandong, 271018, P. R. China

Baohua Xu

qhsun@sdau.edu.cn (QS); bhxu@sdau.edu.cn (BX)

‡ These authors also contributed equally to this work.

Affiliation College of Animal Science and Technology, Shandong Agricultural University, Taian, Shandong, 271018, P. R. China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GL QS BX. Performed the experiments: GL HZ HW. Analyzed the data: HZ XG (fourth author) XG (fifth author). Contributed reagents/materials/analysis tools: QS BX. Wrote the paper: GL HZ.