About the Authors

Xian-Ge Hu

Contributed equally to this work with: Xian-Ge Hu, Hui Liu

Affiliation National Engineering Laboratory for Tree Breeding, Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China

Hui Liu

Contributed equally to this work with: Xian-Ge Hu, Hui Liu

Affiliation National Engineering Laboratory for Tree Breeding, Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China

YuQing Jin

Affiliation National Engineering Laboratory for Tree Breeding, Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China

Yan-Qiang Sun

Affiliation National Engineering Laboratory for Tree Breeding, Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China

Yue Li

Affiliation National Engineering Laboratory for Tree Breeding, Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China

Wei Zhao

Affiliations National Engineering Laboratory for Tree Breeding, Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China, Department of Ecology and Environmental Science, Umeå University, Umeå, Sweden

Yousry A. El-Kassaby

Affiliation Department of Forest and Conservation Sciences, Faculty of Forestry, University of British Columbia, Vancouver, British Columbia, Canada

Xiao-Ru Wang

Affiliations National Engineering Laboratory for Tree Breeding, Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China, Department of Ecology and Environmental Science, Umeå University, Umeå, Sweden

Jian-Feng Mao

jianfeng.mao@bjfu.edu.cn

Affiliation National Engineering Laboratory for Tree Breeding, Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JFM XRW YL. Performed the experiments: XGH YQS YJ ZW HL. Analyzed the data: XGH YQS YJ HL JFM. Contributed reagents/materials/analysis tools: XGH JFM. Wrote the paper: XGH JFM YAE XRW.