About the Authors

Susan N. Bolch

Affiliation Department of Ophthalmology, University of Florida, Gainesville, Florida, United States of America

Donald R. Dugger

Affiliation Department of Ophthalmology, University of Florida, Gainesville, Florida, United States of America

Timothy Chong

Affiliation Department of Ophthalmology, University of Florida, Gainesville, Florida, United States of America

J. Hugh McDowell

Affiliation Department of Ophthalmology, University of Florida, Gainesville, Florida, United States of America

W. Clay Smith

wcsmith@ufl.edu

Affiliation Department of Ophthalmology, University of Florida, Gainesville, Florida, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WCS SNB JHM. Performed the experiments: SNB DRD TC JHM WCS. Analyzed the data: WCS SNB JHM TC. Contributed reagents/materials/analysis tools: DRD WCS JHM. Wrote the paper: WCS SNB.