About the Authors

Elena Arciero

Contributed equally to this work with: Elena Arciero, Simone Andrea Biagini

Affiliations The Wellcome Trust Sanger Institute, Wellcome Genome Campus, Hinxton, CB10 1SA, United Kingdom, Department of Biological, Geological and Environmental Sciences, University of Bologna, 40126, Bologna, Italy

Simone Andrea Biagini

Contributed equally to this work with: Elena Arciero, Simone Andrea Biagini

Current address: Institut de Biologia Evolutiva, CSIC-Universitat Pompeu Fabra, Barcelona, 08003, Spain

Affiliation Department of Biological, Geological and Environmental Sciences, University of Bologna, 40126, Bologna, Italy

Yuan Chen

Affiliation The Wellcome Trust Sanger Institute, Wellcome Genome Campus, Hinxton, CB10 1SA, United Kingdom

Yali Xue

Affiliation The Wellcome Trust Sanger Institute, Wellcome Genome Campus, Hinxton, CB10 1SA, United Kingdom

Donata Luiselli

Affiliation Department of Biological, Geological and Environmental Sciences, University of Bologna, 40126, Bologna, Italy

Chris Tyler-Smith

Affiliation The Wellcome Trust Sanger Institute, Wellcome Genome Campus, Hinxton, CB10 1SA, United Kingdom

Luca Pagani

Affiliations Department of Biological, Geological and Environmental Sciences, University of Bologna, 40126, Bologna, Italy, Division of Biological Anthropology, University of Cambridge, CB2 1QH, Cambridge, United Kingdom

Qasim Ayub

qa1@sanger.ac.uk

Affiliation The Wellcome Trust Sanger Institute, Wellcome Genome Campus, Hinxton, CB10 1SA, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: QA DL LP CT-S. Analyzed the data: EA SAB YC LP QA. Contributed reagents/materials/analysis tools: YC YX QA. Wrote the paper: EA SAB LP QA DL CT-S.