About the Authors

Katherine Gurdziel

Contributed equally to this work with: Katherine Gurdziel, David S. Lorberbaum, Aaron M. Udager, Jane Y. Song

‡ These authors are co-first authors on this work.

Affiliations Department of Cell and Developmental Biology, The University of Michigan, Ann Arbor, MI 48109, United States of America, Department of Computational Medicine and Bioinformatics, The University of Michigan, Ann Arbor, MI 48109, United States of America

David S. Lorberbaum

Contributed equally to this work with: Katherine Gurdziel, David S. Lorberbaum, Aaron M. Udager, Jane Y. Song

‡ These authors are co-first authors on this work.

Affiliations Department of Cell and Developmental Biology, The University of Michigan, Ann Arbor, MI 48109, United States of America, Cellular and Molecular Biology Program, The University of Michigan, Ann Arbor, MI 48109, United States of America

Aaron M. Udager

Contributed equally to this work with: Katherine Gurdziel, David S. Lorberbaum, Aaron M. Udager, Jane Y. Song

‡ These authors are co-first authors on this work.

Affiliation Department of Cell and Developmental Biology, The University of Michigan, Ann Arbor, MI 48109, United States of America

Jane Y. Song

Contributed equally to this work with: Katherine Gurdziel, David S. Lorberbaum, Aaron M. Udager, Jane Y. Song

‡ These authors are co-first authors on this work.

Affiliations Department of Cell and Developmental Biology, The University of Michigan, Ann Arbor, MI 48109, United States of America, Cellular and Molecular Biology Program, The University of Michigan, Ann Arbor, MI 48109, United States of America

Neil Richards

Affiliation Department of Cell and Developmental Biology, The University of Michigan, Ann Arbor, MI 48109, United States of America

David S. Parker

Current address: Department of Biology, Elon University, Elon, NC 27244, United States of America

Affiliation Department of Cell and Developmental Biology, The University of Michigan, Ann Arbor, MI 48109, United States of America

Lisa A. Johnson

Affiliation Department of Cell and Developmental Biology, The University of Michigan, Ann Arbor, MI 48109, United States of America

Benjamin L. Allen

dgumucio@umich.edu (DLG); sbarolo@umich.edu (SB); benallen@umich.edu (BLA)

Affiliation Department of Cell and Developmental Biology, The University of Michigan, Ann Arbor, MI 48109, United States of America

Scott Barolo

dgumucio@umich.edu (DLG); sbarolo@umich.edu (SB); benallen@umich.edu (BLA)

Affiliation Department of Cell and Developmental Biology, The University of Michigan, Ann Arbor, MI 48109, United States of America

Deborah L. Gumucio

dgumucio@umich.edu (DLG); sbarolo@umich.edu (SB); benallen@umich.edu (BLA)

Affiliation Department of Cell and Developmental Biology, The University of Michigan, Ann Arbor, MI 48109, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DLG SB BLA KG DSL JYS AMU. Performed the experiments: KG DSL JYS NR DSP LAJ AMU. Analyzed the data: KG DSL JYS NR DSP LAJ AMU. Wrote the paper: DLG SB BLA KG DSL JYS AMU.