About the Authors

Masaki Horie

Affiliations Division of Medical Science, Faculty of Medicine, University of Tsukuba, Tsukuba, Ibaraki, Japan, Japan Society for the Promotion of Science, Tokyo, Japan

Eiji Warabi

Affiliation Division of Biomedical Sciences, Faculty of Medicine, University of Tsukuba, Tsukuba, Ibaraki, Japan

Shoichi Komine

Affiliation Graduate School of Comprehensive Human Sciences Majors of Medical Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan

Sechang Oh

Affiliations Division of Medical Science, Faculty of Medicine, University of Tsukuba, Tsukuba, Ibaraki, Japan, Japan Society for the Promotion of Science, Tokyo, Japan

Junichi Shoda

shodaj@md.tsukuba.ac.jp

Affiliation Division of Medical Science, Faculty of Medicine, University of Tsukuba, Tsukuba, Ibaraki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MH EW JS. Performed the experiments: MH. Analyzed the data: MH SK SO. Contributed reagents/materials/analysis tools: MH JS. Wrote the paper: MH.