About the Authors

Jeffrey S. Hall

jshall@usgs.gov

Affiliation USGS National Wildlife Health Center, Madison, WI, United States of America, 53711

Robin E. Russell

Affiliation USGS National Wildlife Health Center, Madison, WI, United States of America, 53711

J. Christian Franson

Affiliation USGS National Wildlife Health Center, Madison, WI, United States of America, 53711

Catherine Soos

Affiliation Environment Canada, Science & Technology Branch, Saskatoon, Saskatchewan, Canada, S7N 5B4

Robert J. Dusek

Affiliation USGS National Wildlife Health Center, Madison, WI, United States of America, 53711

R. Bradford Allen

Affiliation Maine Department of Inland Fisheries and Wildlife, Bangor, ME, United States of America, 04401

Sean W. Nashold

Affiliation USGS National Wildlife Health Center, Madison, WI, United States of America, 53711

Joshua L. TeSlaa

Affiliation USGS National Wildlife Health Center, Madison, WI, United States of America, 53711

Jón Eínar Jónsson

Affiliation University of Iceland, Snæfellsnes Research Centre, Stykkishólmur, Iceland 245

Jennifer R. Ballard

Affiliation Southeastern Cooperative Wildlife Disease Study, University of Georgia, Athens, GA, United States of America, 30602

Naomi Jane Harms

Affiliation Department of Veterinary Pathology, Western College of Veterinary Medicine, University of Saskatchewan, Saskatoon, Saskatchewan, Canada, S7N 5B4

Justin D. Brown

Current address: The Pennsylvania State University, University Park, PA, United States of America, 16801

Affiliation Southeastern Cooperative Wildlife Disease Study, University of Georgia, Athens, GA, United States of America, 30602

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JSH RER JCF RBA JDB. Performed the experiments: JSH RER JCF CS RJD SWN JLT JRB JDB. Analyzed the data: JSH RER JDB JRB SWN JLT. Contributed reagents/materials/analysis tools: CS JCF RJD RBA JEJ NJH. Wrote the paper: JSH RER JCF RJD RBA JEJ JDB.