About the Authors

Richard J. Chen

Affiliation Laboratory of Cellular Imaging and Macromolecular Biophysics, National Institute of Biomedical Imaging and Bioengineering, National Institutes of Health, Bethesda, MD, 20892, United States of America

Guofeng Zhang

Affiliation Laboratory of Cellular Imaging and Macromolecular Biophysics, National Institute of Biomedical Imaging and Bioengineering, National Institutes of Health, Bethesda, MD, 20892, United States of America

Susan H. Garfield

Affiliation Experimental Carcinogenesis, National Cancer Institute, National Institutes of Health, Bethesda, MD 20892, United States of America

Yi-Jun Shi

Affiliation NIH Stem Cell Unit, National Institute of Neurological Disorders and Stroke, National Institutes of Health, Bethesda, MD 20892, United States of America

Kevin G. Chen

Affiliation NIH Stem Cell Unit, National Institute of Neurological Disorders and Stroke, National Institutes of Health, Bethesda, MD 20892, United States of America

Pamela G. Robey

Affiliation Craniofacial and Skeletal Diseases Branch, National Institute of Dental and Craniofacial Research, National Institutes of Health, Bethesda, MD, 20892, United States of America

Richard D. Leapman

leapmanr@mail.nih.gov

Affiliation Laboratory of Cellular Imaging and Macromolecular Biophysics, National Institute of Biomedical Imaging and Bioengineering, National Institutes of Health, Bethesda, MD, 20892, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RDL. Performed the experiments: RJC GZ SHG KGC YJS. Analyzed the data: RJC GZ KGC PGR RDL. Wrote the paper: RJC RDL.