About the Authors

Jonathan Lisein

jo.lisein@ulg.ac.be

Affiliations Laboratory of Forest Resources Management, Department of Biosytem Engineering, University of Liège—Gembloux Agro-Bio Tech. 2, Passage des déportés, 5030 Gembloux, Belgium, Ecole nationale des sciences géographiques, 6 et 8 avenue Blaise Pascal, Cité Descartes, Champs-sur-Marne, 77455 Marne la Vallée, France

Adrien Michez

Affiliation Laboratory of Forest Resources Management, Department of Biosytem Engineering, University of Liège—Gembloux Agro-Bio Tech. 2, Passage des déportés, 5030 Gembloux, Belgium

Hugues Claessens

Affiliation Laboratory of Forest Resources Management, Department of Biosytem Engineering, University of Liège—Gembloux Agro-Bio Tech. 2, Passage des déportés, 5030 Gembloux, Belgium

Philippe Lejeune

Affiliation Laboratory of Forest Resources Management, Department of Biosytem Engineering, University of Liège—Gembloux Agro-Bio Tech. 2, Passage des déportés, 5030 Gembloux, Belgium

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JL PL. Performed the experiments: JL AM. Analyzed the data: JL AM HC PL. Contributed reagents/materials/analysis tools: PL HC. Wrote the paper: JL HC AM.