About the Authors

Nami Goto-Yamamoto

gotoh_n@nrib.go.jp (NG); sean.myles@dal.ca (SM)

Affiliation National Research Institute of Brewing, Higashi-Hiroshima, Japan

Jason Sawler

Affiliations Faculty of Agriculture, Dalhousie University, Truro, Nova Scotia, Canada, Anandia Labs, 2259 Lower Mall, Vancouver, British Columbia, Canada

Sean Myles

gotoh_n@nrib.go.jp (NG); sean.myles@dal.ca (SM)

Affiliation Faculty of Agriculture, Dalhousie University, Truro, Nova Scotia, Canada

Competing Interests

The authors have the following interests: Co-author Jason Sawler is employed by Anandia Labs. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: NG SM. Performed the experiments: NG. Analyzed the data: NG JS. Contributed reagents/materials/analysis tools: NG SM. Wrote the paper: NG JS SM.