About the Authors

Guiying Tang

Affiliation Bio-Tech Research Centre, Shandong Academy of Agricultural Sciences / Shandong Provincial Key Laboratory of Crop Genetic Improvement, Ecology and Physiology, Jinan, 250100, China

Pingli Xu

Affiliation Bio-Tech Research Centre, Shandong Academy of Agricultural Sciences / Shandong Provincial Key Laboratory of Crop Genetic Improvement, Ecology and Physiology, Jinan, 250100, China

Wei Liu

Affiliation College of Agriculture, Shandong University, Jinan, 250100, China

Zhanji Liu

Affiliation Bio-Tech Research Centre, Shandong Academy of Agricultural Sciences / Shandong Provincial Key Laboratory of Crop Genetic Improvement, Ecology and Physiology, Jinan, 250100, China

Lei Shan

Shlei1025@sina.com

Affiliations Bio-Tech Research Centre, Shandong Academy of Agricultural Sciences / Shandong Provincial Key Laboratory of Crop Genetic Improvement, Ecology and Physiology, Jinan, 250100, China, College of Agriculture, Shandong University, Jinan, 250100, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LS. Performed the experiments: GYT PLX WL. Analyzed the data: LS ZJL. Contributed reagents/materials/analysis tools: GYT PLX WL ZJL. Wrote the paper: LS.