About the Authors

Tiejun Zhang

Contributed equally to this work with: Tiejun Zhang, Long-Xi Yu, Ping Zheng

Current address: Institute of Animal Science, Chinese Academy of Agricultural Sciences, Beijing, China

Affiliation Plant and Germplasm Introduction and Testing Research, United States Department of Agriculture-Agricultural Research Service, Prosser, Washington, United States of America

Long-Xi Yu

Contributed equally to this work with: Tiejun Zhang, Long-Xi Yu, Ping Zheng

longxi.yu@ars.usda.gov

Affiliation Plant and Germplasm Introduction and Testing Research, United States Department of Agriculture-Agricultural Research Service, Prosser, Washington, United States of America

Ping Zheng

Contributed equally to this work with: Tiejun Zhang, Long-Xi Yu, Ping Zheng

Affiliation Department of Horticulture, Washington State University, Pullman, Washington, United States of America

Yajun Li

Affiliation Plant and Germplasm Introduction and Testing Research, United States Department of Agriculture-Agricultural Research Service, Prosser, Washington, United States of America

Martha Rivera

Affiliation Plant and Germplasm Introduction and Testing Research, United States Department of Agriculture-Agricultural Research Service, Prosser, Washington, United States of America

Dorrie Main

Affiliation Department of Horticulture, Washington State University, Pullman, Washington, United States of America

Stephanie L. Greene

Affiliation National Center for Genetic Resource Preservation, United States Department of Agriculture-Agricultural Research Service, Fort Collins, Colorado, United States of America

Competing Interests

This work was partly supported by a grant from the National Alfalfa and Forage Alliance. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: L-XY. Performed the experiments: TZ YL MR. Analyzed the data: PZ. Wrote the paper: SG DM.