About the Authors

Sarah L. Shafer

sshafer@usgs.gov

Affiliation U. S. Geological Survey, Corvallis, Oregon, United States of America

Patrick J. Bartlein

Affiliation Department of Geography, University of Oregon, Eugene, Oregon, United States of America

Elizabeth M. Gray

Affiliation The Nature Conservancy, Maryland/DC, Bethesda, Maryland, United States of America

Richard T. Pelltier

Affiliation U. S. Geological Survey, Denver, Colorado, United States of America

Competing Interests

The authors have read the journal's policy have the following competing interests: SLS and RTP are employed by the U.S. Geological Survey, but in a different part of the organization than the funders of this work. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SLS EMG. Wrote the paper: SLS PJB EMG RTP. Created and analyzed data: SLS PJB RTP.