About the Authors

Zhong Chen

Contributed equally to this work with: Zhong Chen, Kai Gao

Affiliations National Engineering Laboratory for Tree Breeding, College of Biological Sciences and Biotechnology, Beijing Forestry University, Beijing, China, Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants of the Ministry of Education, College of Biological Sciences and Biotechnology, Beijing Forestry University, Beijing, China

Kai Gao

Contributed equally to this work with: Zhong Chen, Kai Gao

Affiliations National Engineering Laboratory for Tree Breeding, College of Biological Sciences and Biotechnology, Beijing Forestry University, Beijing, China, Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants of the Ministry of Education, College of Biological Sciences and Biotechnology, Beijing Forestry University, Beijing, China

Xiaoxing Su

Affiliation Beijing Berry Genomics Company Limited, Beijing, China

Pian Rao

Affiliations National Engineering Laboratory for Tree Breeding, College of Biological Sciences and Biotechnology, Beijing Forestry University, Beijing, China, Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants of the Ministry of Education, College of Biological Sciences and Biotechnology, Beijing Forestry University, Beijing, China

Xinmin An

anxinmin@bjfu.edu.cn

Affiliations National Engineering Laboratory for Tree Breeding, College of Biological Sciences and Biotechnology, Beijing Forestry University, Beijing, China, Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants of the Ministry of Education, College of Biological Sciences and Biotechnology, Beijing Forestry University, Beijing, China

Competing Interests

XS is an employee of Beijing Berry Genomics Company Limited, a funder of this study. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ZC XA. Performed the experiments: ZC PR KG. Analyzed the data: ZC XS PR. Contributed reagents/materials/analysis tools: ZC XS PR KG. Wrote the paper: ZC.