About the Authors

Andrew S. Flies

andyflies@gmail.com

Affiliations Menzies Research Institute Tasmania, University of Tasmania, Hobart, TAS, Australia, Department of Zoology, Michigan State University, East Lansing, MI, United States of America, Interdisciplinary program in Ecology, Evolutionary Biology and Behavior, Michigan State University, East Lansing, MI, United States of America, Department of Pharmacy and Medical Sciences, University of South Australia, Adelaide, South Australia, Australia

Linda S. Mansfield

Affiliation Department of Microbiology and Molecular Genetics, National Food Safety and Toxicology Center, Michigan State University, East Lansing, MI, United States of America

Chris K. Grant

Affiliation Custom Monoclonals International Corp, West Sacramento, CA, United States of America

Mary L. Weldele

Affiliation Department of Psychology, University of California, Berkeley, CA, United States of America

Kay E. Holekamp

Affiliations Department of Zoology, Michigan State University, East Lansing, MI, United States of America, Interdisciplinary program in Ecology, Evolutionary Biology and Behavior, Michigan State University, East Lansing, MI, United States of America

Competing Interests

The authors have the following competing interests. Chris K. Grant is president of Custom Monoclonals International, which contributed antibodies for this research. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: ASF LSM CKG KEH. Performed the experiments: ASF MLW. Analyzed the data: ASF LSM KEH. Contributed reagents/materials/analysis tools: ASF LSM CKG KEH MLW. Wrote the paper: ASF LSM CKG KEH. Submitted the manuscript: ASF.