About the Authors

Efstathios S. Giotis

e.giotis@imperial.ac.uk

Current address: Section of Virology, Imperial College London, Faculty of Medicine, London, United Kingdom

Affiliations Agri-Food and Biosciences Institute, Belfast, United Kingdom, Queen’s University Belfast, Belfast, United Kingdom, The Roslin Institute and R(D)SVS, University of Edinburgh, Edinburgh, United Kingdom

Lisa Rothwell

Current address: The Roslin Institute and R(D)SVS, University of Edinburgh, Edinburgh, United Kingdom

Affiliation Institute for Animal Health, Compton, United Kingdom

Alistair Scott

Affiliation Queen’s University Belfast, Belfast, United Kingdom

Tuanjun Hu

Current address: The Roslin Institute and R(D)SVS, University of Edinburgh, Edinburgh, United Kingdom

Affiliation Institute for Animal Health, Compton, United Kingdom

Richard Talbot

Affiliation The Roslin Institute and R(D)SVS, University of Edinburgh, Edinburgh, United Kingdom

Daniel Todd

Affiliation Agri-Food and Biosciences Institute, Belfast, United Kingdom

David W. Burt

Affiliation The Roslin Institute and R(D)SVS, University of Edinburgh, Edinburgh, United Kingdom

Elizabeth J. Glass

Affiliation The Roslin Institute and R(D)SVS, University of Edinburgh, Edinburgh, United Kingdom

Pete Kaiser

Current address: The Roslin Institute and R(D)SVS, University of Edinburgh, Edinburgh, United Kingdom

Affiliation Institute for Animal Health, Compton, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DT DB EJG PK. Performed the experiments: ESG AS DT TH LR. Analyzed the data: ESG RT TH LR DB. Contributed reagents/materials/analysis tools: RT DT DB EJG PK. Wrote the paper: ESG DT DB EJG PK.