About the Authors

Dong-He Li

Contributed equally to this work with: Dong-He Li, Wei Shi

Affiliations Key Laboratory of Tropical Marine Bio-resources and Ecology and Guangdong Provincial Key Laboratory of Applied Marine Biology, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, 510301, China, South China Sea Bio-Resource Exploitation and Utilization Collaborative Innovation Center, Guangzhou, 510301, China, University of Chinese Academy of Sciences, Beijing, 100049, China

Wei Shi

Contributed equally to this work with: Dong-He Li, Wei Shi

shiwei@scsio.ac.cn (WS); xykong@scsio.ac.cn (XYK)

Affiliations Key Laboratory of Tropical Marine Bio-resources and Ecology and Guangdong Provincial Key Laboratory of Applied Marine Biology, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, 510301, China, South China Sea Bio-Resource Exploitation and Utilization Collaborative Innovation Center, Guangzhou, 510301, China

Thomas A. Munroe

Affiliation National Systematics Laboratory NMFS/NOAA, Post Office Box 37012, Smithsonian Institution NHB, WC 60, MRC-153, Washington, D.C., 20013–7012, United States of America

Li Gong

Affiliations Key Laboratory of Tropical Marine Bio-resources and Ecology and Guangdong Provincial Key Laboratory of Applied Marine Biology, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, 510301, China, South China Sea Bio-Resource Exploitation and Utilization Collaborative Innovation Center, Guangzhou, 510301, China, University of Chinese Academy of Sciences, Beijing, 100049, China

Xiao-Yu Kong

shiwei@scsio.ac.cn (WS); xykong@scsio.ac.cn (XYK)

Affiliation Key Laboratory of Tropical Marine Bio-resources and Ecology and Guangdong Provincial Key Laboratory of Applied Marine Biology, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, 510301, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XYK WS. Performed the experiments: DHL LG. Analyzed the data: WS DHL. Contributed reagents/materials/analysis tools: WS. Wrote the paper: WS. Revised the manuscript critically for important intellectual content: TAM.