About the Authors

Gyoungju Nah

Affiliation Department of Plant Science, Research Institute of Agriculture and Life Sciences, College of Agriculture and Life Sciences, Seoul National University, Seoul 151–921, Korea

Ji-Hoon Im

Affiliation Department of Plant Science, Research Institute of Agriculture and Life Sciences, College of Agriculture and Life Sciences, Seoul National University, Seoul 151–921, Korea

Jin-Won Kim

Affiliation Department of Plant Science, Research Institute of Agriculture and Life Sciences, College of Agriculture and Life Sciences, Seoul National University, Seoul 151–921, Korea

Hae-Rim Park

Affiliation Department of Plant Science, Research Institute of Agriculture and Life Sciences, College of Agriculture and Life Sciences, Seoul National University, Seoul 151–921, Korea

Min-Jung Yook

Affiliation Department of Plant Science, Research Institute of Agriculture and Life Sciences, College of Agriculture and Life Sciences, Seoul National University, Seoul 151–921, Korea

Tae-Jin Yang

Affiliation Department of Plant Science, Research Institute of Agriculture and Life Sciences, College of Agriculture and Life Sciences, Seoul National University, Seoul 151–921, Korea

Albert J. Fischer

Affiliation Department of Plant Sciences, College of Agricultural and Environmental Sciences, University of California Davis, Davis, CA 95616, United States of America

Do-Soon Kim

dosoonkim@snu.ac.kr

Affiliation Department of Plant Science, Research Institute of Agriculture and Life Sciences, College of Agriculture and Life Sciences, Seoul National University, Seoul 151–921, Korea

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DK GN. Performed the experiments: JI HP. Analyzed the data: GN. Contributed reagents/materials/analysis tools: JK. Wrote the paper: GN DK MY AF TY.