About the Authors

Gary A. Clawson

gac4@psu.edu

Affiliation Department of Pathology and Gittlen Cancer Research Laboratories, Hershey Medical Center, Pennsylvania State University, Hershey, Pennsylvania, United States of America

Gail L. Matters

Affiliation Department of Biochemistry & Molecular Biology, Hershey Medical Center, Pennsylvania State University, Hershey, Pennsylvania, United States of America

Ping Xin

Affiliation Department of Pathology and Gittlen Cancer Research Laboratories, Hershey Medical Center, Pennsylvania State University, Hershey, Pennsylvania, United States of America

Yuka Imamura-Kawasawa

Affiliation Department of Pharmacology and the Institute for Personalized Medicine, Hershey Medical Center, Pennsylvania State University, Hershey, Pennsylvania, United States of America

Zhen Du

Affiliation Department of Pathology and Gittlen Cancer Research Laboratories, Hershey Medical Center, Pennsylvania State University, Hershey, Pennsylvania, United States of America

Diane M. Thiboutot

Affiliation Department of Dermatology, Division of Health Science Research, Hershey Medical Center, Pennsylvania State University, Hershey, Pennsylvania, United States of America

Klaus F. Helm

Affiliation Department of Dermatopathology, Hershey Medical Center, Pennsylvania State University, Hershey, Pennsylvania, United States of America

Rogerio I. Neves

Affiliation Department of Surgery and the Melanoma Center, Hershey Medical Center, Pennsylvania State University, Hershey, Pennsylvania, United States of America

Thomas Abraham

Affiliation Department of Neural and Behavioral Science and the Microscopy Imaging Facility, Hershey Medical Center, Pennsylvania State University, Hershey, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GAC GLM DMT RIN. Performed the experiments: PX YIK KFH TA ZD. Analyzed the data: GAC PX YIK KFH TA. Contributed reagents/materials/analysis tools: KFH TA RIN. Wrote the paper: GAC GLM PX YIK DMT KFH RIN TA.