About the Authors

Joanne A. M. van der Borg

joanne.vanderborg@wur.nl

Affiliation Wageningen University Behavioural Ecology Group, Department of Animal Sciences, P.O. Box 338, 6700 AH, Wageningen, The Netherlands

Matthijs B. H. Schilder

Affiliation Utrecht University Faculty of Veterinary Medicine, Department of Animals in Science & Society, P.O. Box 80166, 3508 TD, Utrecht, The Netherlands

Claudia M. Vinke

Affiliation Utrecht University Faculty of Veterinary Medicine, Department of Animals in Science & Society, P.O. Box 80166, 3508 TD, Utrecht, The Netherlands

Han de Vries

Affiliation Utrecht University Animal Ecology Group, Department of Biology, Padualaan 8, f3584 CH, Utrecht, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JvdB. Performed the experiments: JvdB. Analyzed the data: JvdB HdV MS CV. Wrote the paper: JvdB MS CV HdV.