About the Authors

Matthew Nahorniak

matt@southforkresearch.org

Affiliation South Fork Research, Inc., North Bend, Washington, United States of America

David P. Larsen

Contributed equally to this work with: David P. Larsen, Carol Volk, Chris E. Jordan

Affiliation Pacific States Marine Fisheries Commission, Corvallis, Oregon, United States of America

Carol Volk

Contributed equally to this work with: David P. Larsen, Carol Volk, Chris E. Jordan

Affiliation South Fork Research, Inc., North Bend, Washington, United States of America

Chris E. Jordan

Contributed equally to this work with: David P. Larsen, Carol Volk, Chris E. Jordan

Affiliation Northwest Fisheries Science Center, NOAA-Fisheries, Seattle, Washington, United States of America

Competing Interests

Two of the authors (MN, CV) are employed by South Fork Research, Inc. South Fork research receives funding to support this research via contracts with the Bonneville Power Administration. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MN DPL CV CEJ. Performed the experiments: MN. Analyzed the data: MN DPL CV CEJ. Contributed reagents/materials/analysis tools: MN DPL CV CEJ. Wrote the paper: MN DPL CV CEJ.