About the Authors

David C. Evans

d.evans@utoronto.ca

Affiliations Department of Natural History, Royal Ontario Museum, 100 Queen’s Park, Toronto, Ontario, M5S 2C6 Canada, Department of Ecology and Evolutionary Biology, University of Toronto, 25 Willcocks Street, Toronto, Ontario, M5S 3B2 Canada

Michael J. Ryan

Affiliation Department of Vertebrate Paleontology, Cleveland Museum of Natural History, 1 Wade Oval Drive, University Circle, Cleveland, Ohio 44106, United States of America

Competing Interests

DCE is currently serving on the editorial board of PLOS ONE as an Academic Editor. MJR has no competing interests. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: DCE MJR. Performed the experiments: DCE MJR. Analyzed the data: DCE MJR. Contributed reagents/materials/analysis tools: DCE MJR. Wrote the paper: DCE MJR.