About the Authors

Eui-Soo Kim

Affiliations Animal Genomics & Improvement Laboratory, Beltsville Agricultural Research Center, Agricultural Research Service, United States Department of Agriculture, Beltsville, Maryland, United States of America, Department of Animal Science, Iowa State University, Ames, Iowa, United States of America

Tad S. Sonstegard

Affiliation Animal Genomics & Improvement Laboratory, Beltsville Agricultural Research Center, Agricultural Research Service, United States Department of Agriculture, Beltsville, Maryland, United States of America

Curtis P. Van Tassell

Affiliation Animal Genomics & Improvement Laboratory, Beltsville Agricultural Research Center, Agricultural Research Service, United States Department of Agriculture, Beltsville, Maryland, United States of America

George Wiggans

Affiliation Animal Genomics & Improvement Laboratory, Beltsville Agricultural Research Center, Agricultural Research Service, United States Department of Agriculture, Beltsville, Maryland, United States of America

Max F. Rothschild

mfrothsc@iastate.edu

Affiliation Department of Animal Science, Iowa State University, Ames, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ESK CPVT TSS. Performed the experiments: TSS. Analyzed the data: ESK GW. Contributed reagents/materials/analysis tools: TSS MFR CPVT. Wrote the paper: ESK MFR TSS CPVT GW.