About the Authors

Alevtina Gall

Affiliations Molecular and Cellular Biology Graduate Program, University of Washington, Seattle, Washington, United States of America, Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Jutta Fero

Affiliation Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Connor McCoy

Affiliations Divisions of Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Program in Computational Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Brian C. Claywell

Affiliations Divisions of Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Program in Computational Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Carissa A. Sanchez

Affiliation Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Patricia L. Blount

Affiliations Divisions of Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Department of Medicine, University of Washington School of Medicine, Seattle, Washington, United States of America

Xiaohong Li

Affiliations Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Divisions of Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Thomas L. Vaughan

Affiliations Divisions of Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Department of Epidemiology, University of Washington, Seattle, Washington, United States of America

Frederick A. Matsen

Affiliations Divisions of Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Program in Computational Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Brian J. Reid

Affiliations Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Divisions of Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Department of Medicine, University of Washington School of Medicine, Seattle, Washington, United States of America, Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Nina R. Salama

nsalama@fhcrc.org

Affiliations Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Department of Microbiology, University of Washington School of Medicine, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NRS BJR TLV. Performed the experiments: AG JF CAS PLB NRS. Analyzed the data: AG CM BCC XL FAM. Contributed reagents/materials/analysis tools: CM BCC TLV FAM. Wrote the paper: AG CAS PLB XL TLV FAM BJR NRS.