About the Authors

David B. Nicholson

david.nicholson@nhm.ac.uk

Current address: Department of Earth Sciences, The Natural History Museum, London, United Kingdom

Affiliations Department of Biology, University of York, York, United Kingdom, Department of Natural Sciences, National Museum of Scotland, Edinburgh, United Kingdom

Peter J. Mayhew

Affiliation Department of Biology, University of York, York, United Kingdom

Andrew J. Ross

Affiliation Department of Natural Sciences, National Museum of Scotland, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AJR PJM DBN. Analyzed the data: DBN PJM AJR. Wrote the paper: DBN PJM AJR.