About the Authors

Tom N. McNeilly

Tom.McNeilly@moredun.ac.uk

Affiliation Moredun Research Institute, Edinburgh, United Kingdom

Mairi C. Mitchell

Affiliation Moredun Research Institute, Edinburgh, United Kingdom

Alexander Corbishley

Affiliations Moredun Research Institute, Edinburgh, United Kingdom, Division of Immunity and Infection, The Roslin Institute, University of Edinburgh, Edinburgh, United Kingdom

Mintu Nath

Affiliation Biomathematics and Statistics Scotland, Edinburgh, United Kingdom

Hannah Simmonds

Affiliation Division of Immunity and Infection, The Roslin Institute, University of Edinburgh, Edinburgh, United Kingdom

Sean P. McAteer

Affiliation Division of Immunity and Infection, The Roslin Institute, University of Edinburgh, Edinburgh, United Kingdom

Arvind Mahajan

Affiliation Division of Immunity and Infection, The Roslin Institute, University of Edinburgh, Edinburgh, United Kingdom

J. Christopher Low

Affiliation Division of Immunity and Infection, The Roslin Institute, University of Edinburgh, Edinburgh, United Kingdom

David G. E. Smith

Affiliations Moredun Research Institute, Edinburgh, United Kingdom, Institute of Infection, Immunity & Inflammation, University of Glasgow, Glasgow, United Kingdom

John F. Huntley

Affiliation Moredun Research Institute, Edinburgh, United Kingdom

David L. Gally

Affiliation Division of Immunity and Infection, The Roslin Institute, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TNM AM JCL DGES JFH DLG. Performed the experiments: TNM MCM AC HS SPM. Analyzed the data: TNM MN. Contributed reagents/materials/analysis tools: TNM MCM AC HS SPM. Wrote the paper: TNM MCM AC MN HS SPM AM JCL DGES JFH DLG.