About the Authors

Courtney M. Tate

Contributed equally to this work with: Courtney M. Tate, Jacquelyn Mc Entire

Affiliation Discovery Research, Eli Lilly and Company, Indianapolis, United States of America

Jacquelyn Mc Entire

Contributed equally to this work with: Courtney M. Tate, Jacquelyn Mc Entire

Affiliation Discovery Research, Eli Lilly and Company, Indianapolis, United States of America

Roberto Pallini

Affiliation Department of Neurosurgery, Università Cattolica del Sacro Cuore, Rome, Italy

Eliza Vakana

Affiliation Discovery Research, Eli Lilly and Company, Indianapolis, United States of America

Lisa Wyss

Affiliation Discovery Research, Eli Lilly and Company, Indianapolis, United States of America

Wayne Blosser

Affiliation Discovery Research, Eli Lilly and Company, Indianapolis, United States of America

Lucia Ricci-Vitiani

Affiliation Department of Hematology, Oncology and Molecular Medicine, Istituto Superiore di Sanità, Rome, Italy

Quintino Giorgio D’Alessandris

Affiliation Department of Neurosurgery, Università Cattolica del Sacro Cuore, Rome, Italy

Liliana Morgante

Affiliation Institute of Human Anatomy, Università Cattolica del Sacro Cuore, Rome, Italy

Stefano Giannetti

Affiliation Institute of Human Anatomy, Università Cattolica del Sacro Cuore, Rome, Italy

Luigi Maria Larocca

Affiliation Institute of Pathology, Università Cattolica del Sacro Cuore, Rome, Italy

Matilde Todaro

Affiliation Surgical and Oncological Sciences, University of Palermo, Palermo, Italy

Antonina Benfante

Affiliation Surgical and Oncological Sciences, University of Palermo, Palermo, Italy

Maria Luisa Colorito

Affiliation Surgical and Oncological Sciences, University of Palermo, Palermo, Italy

Giorgio Stassi

Affiliation Surgical and Oncological Sciences, University of Palermo, Palermo, Italy

Ruggero De Maria

Affiliation Regina Elena National Cancer Institute, Rome, Italy

Scott Rowlinson

Affiliation Discovery Research, Eli Lilly and Company, Indianapolis, United States of America

Louis Stancato

l.stancato@lilly.com (LS)

Affiliation Discovery Research, Eli Lilly and Company, Indianapolis, United States of America

Competing Interests

Authors affiliated with Eli Lilly and Company have Eli Lilly and Company shares, received via 401(k) and bonus plans, and have an affiliation to the commercial funders of this research (Eli Lilly). This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: CMT JM RP AB RD SR LRV LS. Performed the experiments: CMT JM RP EV LW WB QGD LM SG LML GS AB MLC MT. Analyzed the data: CMT JM RP EV LW QGD LM SG LML GS AB MLC MT. Wrote the paper: CMT JM RP SR LS.